John Lilly
InvestorPartner at Greylock Partners Β· Advising Partner at Gigascale Capital Β· Board Member (Partner, Greylock Partners) at Dwnld Β· Director; Lead Independent Director (Greylock board partner) at Figma
San Francisco Bay Area
About
John Lilly is an Advising Partner at Gigascale Capital with roughly 30 years of experience as a founder, CEO, and investor. He previously spent 8 years as a General Partner at Greylock and was CEO of Mozilla, where he oversaw Firefox's growth. Earlier he co-founded the enterprise security company Reactivity, later acquired by Cisco, and held roles at Apple and Trilogy Software. He is also a Lecturer at Stanford's Graduate School of Business and holds BS and MS degrees from Stanford.
Likely emailsΒ· guessed from name + firm domain, not firm-published
- jβ’β’β’@greylock.comreveal100% confident
- jβ’β’β’@greylock.comreveal90% confident
Invests in
CybersecurityEnterprise Software
Board Member (Partner, Greylock Partners) at Dwnld
Intro paths
In turn, John can intro founders to Dwnld's investors below.
Dwnld's investors Β· 3
Co-founders at Dwnld
Director; Lead Independent Director (Greylock board partner) at Figma
Intro paths
In turn, John can intro founders to Figma's investors below.
Figma's investors Β· 33
Co-founders at Figma

Experience
- Gigascale CapitalAdvising Partnercurrent2026 β present
- FigmaBoard Membercurrent2014 β present
- DuolingoBoard Membercurrent2021 β present
- VotingWorksBoard Membercurrent2018 β present
- NuroBoard Membercurrent2019 β present
- Greylock PartnersVenture Partnercurrent2019 β present
- Greylock PartnersPartner2011 β 2019
- Stanford UniversityConsulting Assistant Professor & Lecturercurrent2008 β present
- Code for AmericaChair, Board Of Directors2017 β 2025
- Code for AmericaBoard Member2012 β 2025
- ClockwiseBoard Member2018 β 2025
- CaffeineBoard Member2016 β 2024
- MozillaBoard Member2006 β 2014
- MozillaCEO2008 β 2010
- MozillaChief Operating Officer & Board Member2005 β 2007
- TripItBoard of Directors2008 β 2010
- Sunnyvale Public LibraryMember, Board of Trustees2005 β 2009
- Open Source Applications FoundationMember, Board of Directors2001 β 2008
- Reactivity (acquired by Cisco)CTO, VP Products, Board of Directors & Founder2001 β 2004
- Reactivity (acquired by Cisco)CEO, Board of Directors & Founder1998 β 2000
- CenterRunMember, Board of Directors2000 β 2003
- Apple ComputerSenior Scientist1997
- Trilogy SoftwareDirector of Design1995 β 1996
Skills & more
+12 more
Product ManagementBusiness DevelopmentUser Interface DesignScalabilityStart-upsUser InterfaceStrategic PartnershipsUser-centered DesignWeb DevelopmentEntrepreneurshipMobile DevicesLeadershipFrom John's professional profile.
Deals Β· 2
FigmaBoard seat
via Greylock Partners Β· filed Dec 2015 Β· $14M raise
DwnldBoard seat
via Greylock Partners Β· filed Sep 2015 Β· $17.1M raise
Attribution from SEC filings, press coverage, and firm rosters.
Other disclosed investments
Led
DropboxInstagramQuipTumblr
Board seats
DuolingoNuroQuip
Sources: the firm's website Β· SEC Form D filings.
Colleagues at Greylock Partners
Network2
Investors connected to John in public investor directories β a warm-intro map.
Frequently asked questions
- Who is John Lilly?
- John Lilly is an Advising Partner at Gigascale Capital with roughly 30 years of experience as a founder, CEO, and investor. He previously spent 8 years as a General Partner at Greylock and was CEO of Mozilla, where he oversaw Firefox's growth. Earlier he co-founded the enterprise security company Reactivity, later acquired by Cisco, and held roles at Apple and Trilogy Software. He is also a Lecturer at Stanford's Graduate School of Business and holds BS and MS degrees from Stanford.
- Where does John Lilly work?
- John Lilly is Partner at Greylock Partners and Board Member (Partner, Greylock Partners) at Dwnld and Director; Lead Independent Director (Greylock board partner) at Figma.
- What does John Lilly invest in?
- John Lilly invests in Cybersecurity, Enterprise Software.
- What boards does John Lilly sit on?
- John Lilly holds tracked board seats at Duolingo, Dwnld, Figma, Figma, Nuro, Quip.
- What companies has John Lilly invested in?
- Investments tracked here include Dropbox, Figma, Instagram, Quip, Tumblr.
- Where did John Lilly go to school?
- John Lilly studied at Stanford University (MS, Computer Science); Stanford University (BS, Computer Systems Engineering).
- Does John Lilly lead rounds?
- Yes β John Lilly has led 5 tracked investments.
Same school
Also attended Stanford University2,000
+ 1,994 more
Shared employers
Also worked at Nuro12
+ 6 more
Also worked at Clockwise5
Also worked at Mozilla42
+ 36 more











































.jpg)








