John Lilly
InvestorAdvising Partner at Gigascale Capital · Consulting Assistant Professor & Lecturer at Stanford University · Venture Partner at Greylock Partners · Board Member / Board Partner (Greylock) at Clockwise · Board Member; Venture Partner, Greylock Partners; Advising Partner, Gigascale Capital at Nuro · +1 more
San Francisco Bay Area
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You →Chad · together at Trilogy Software (1995–1996) →John
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About
John Lilly is an Advising Partner at Gigascale Capital with roughly 30 years of experience as a founder, CEO, and investor. He previously spent 8 years as a General Partner at Greylock and was CEO of Mozilla, where he oversaw Firefox's growth. Earlier he co-founded the enterprise security company Reactivity, later acquired by Cisco, and held roles at Apple and Trilogy Software. He is also a Lecturer at Stanford's Graduate School of Business and holds BS and MS degrees from Stanford.
Likely emails· guessed from name + firm domain, not firm-published
- j•••@greylock.comreveal100% confident
- j•••@greylock.comreveal90% confident
Invests in
CybersecurityEnterprise Software
Board Member / Board Partner (Greylock) at Clockwise
Intro paths
In turn, John can intro founders to Clockwise's investors below.
Clockwise's investors · 10
Co-founders at Clockwise
Board Member; Venture Partner, Greylock Partners; Advising Partner, Gigascale Capital at Nuro
Intro paths
In turn, John can intro founders to Nuro's investors below.
Nuro's investors · 22
Co-founders at Nuro


Board Member (Partner, Greylock Partners) at Dwnld
Intro paths
In turn, John can intro founders to Dwnld's investors below.
Dwnld's investors · 4
Co-founders at Dwnld


Experience
- Gigascale CapitalAdvising Partnercurrent2026 — present
- FigmaBoard Membercurrent2014 — present
- DuolingoBoard Membercurrent2021 — present
- VotingWorksBoard Membercurrent2018 — present
- NuroBoard Membercurrent2019 — present
- Greylock PartnersVenture Partnercurrent2019 — present
- Greylock PartnersPartner2011 — 2019
- Stanford UniversityConsulting Assistant Professor & Lecturercurrent2008 — present
- Code for AmericaChair, Board Of Directors2017 — 2025
- Code for AmericaBoard Member2012 — 2025
- ClockwiseBoard Member2018 — 2025
- CaffeineBoard Member2016 — 2024
- MozillaBoard Member2006 — 2014
- MozillaCEO2008 — 2010
- MozillaChief Operating Officer & Board Member2005 — 2007
- TripItBoard of Directors2008 — 2010
- Sunnyvale Public LibraryMember, Board of Trustees2005 — 2009
- Open Source Applications FoundationMember, Board of Directors2001 — 2008
- Reactivity (acquired by Cisco)CTO, VP Products, Board of Directors & Founder2001 — 2004
- Reactivity (acquired by Cisco)CEO, Board of Directors & Founder1998 — 2000
- CenterRunMember, Board of Directors2000 — 2003
- Apple ComputerSenior Scientist1997
- Trilogy SoftwareDirector of Design1995 — 1996
Skills & more
+8 more
Business DevelopmentUser Interface DesignScalabilityStart-upsUser InterfaceStrategic PartnershipsSaaSUser-centered DesignFrom John's professional profile.
Deals · 2
FigmaBoard seat
via Greylock Partners · filed Dec 2015 · $14M raise
DwnldBoard seat
via Greylock Partners · filed Sep 2015 · $17.1M raise
Attribution from SEC filings, press coverage, and firm rosters.
Other disclosed investments
Led
DropboxInstagramQuipTumblr
Board seats
DuolingoNuroQuip
Sources: the firm's website · SEC Form D filings · curated public sources.
Colleagues at Gigascale Capital
Network2
Investors connected to John in public investor directories — a warm-intro map.
Frequently asked questions
- Who is John Lilly?
- John Lilly is an Advising Partner at Gigascale Capital with roughly 30 years of experience as a founder, CEO, and investor. He previously spent 8 years as a General Partner at Greylock and was CEO of Mozilla, where he oversaw Firefox's growth. Earlier he co-founded the enterprise security company Reactivity, later acquired by Cisco, and held roles at Apple and Trilogy Software. He is also a Lecturer at Stanford's Graduate School of Business and holds BS and MS degrees from Stanford.
- Where does John Lilly work?
- John Lilly is Advising Partner at Gigascale Capital and Board Member / Board Partner (Greylock) at Clockwise and Board Member; Venture Partner, Greylock Partners; Advising Partner, Gigascale Capital at Nuro and Board Member (Partner, Greylock Partners) at Dwnld.
- What does John Lilly invest in?
- John Lilly invests in Cybersecurity, Enterprise Software.
- What boards does John Lilly sit on?
- John Lilly holds tracked board seats at Duolingo, Dwnld, Figma, Figma, Nuro, Quip.
- What companies has John Lilly invested in?
- Investments tracked here include Dropbox, Figma, Instagram, Quip, Tumblr, Airtop, Apprente, Arcol, Aside, Basedash, Census, Citrus Lane, and 35 more.
- Where did John Lilly go to school?
- John Lilly studied at Stanford University (MS, Computer Science); Stanford University (BS, Computer Systems Engineering).
- Does John Lilly lead rounds?
- Yes — John Lilly has led 5 tracked investments.
Founder mafias
Stanford University mafia2,274
Also came through, then founded or led a company.
Greylock Partners mafia32
Also came through, then founded or led a company.
Figma mafia27
Also came through, then founded or led a company.
Nuro mafia14
Also came through, then founded or led a company.
Duolingo mafia12
Also came through, then founded or led a company.
Same school
Also attended Stanford University2,000
+ 1,994 more
Shared employers
Also worked at Nuro18
+ 12 more
Also worked at Clockwise12
+ 6 more
Also worked at Mozilla81
+ 75 more



























































































