Fundraising Fox

John Lilly

Investor

Partner at Greylock Partners Β· Advising Partner at Gigascale Capital Β· Board Member (Partner, Greylock Partners) at Dwnld Β· Director; Lead Independent Director (Greylock board partner) at Figma

San Francisco Bay Area

About

John Lilly is an Advising Partner at Gigascale Capital with roughly 30 years of experience as a founder, CEO, and investor. He previously spent 8 years as a General Partner at Greylock and was CEO of Mozilla, where he oversaw Firefox's growth. Earlier he co-founded the enterprise security company Reactivity, later acquired by Cisco, and held roles at Apple and Trilogy Software. He is also a Lecturer at Stanford's Graduate School of Business and holds BS and MS degrees from Stanford.

Likely emailsΒ· guessed from name + firm domain, not firm-published

Invests in

CybersecurityEnterprise Software

Board Member (Partner, Greylock Partners) at Dwnld

Intro paths

β†’

In turn, John can intro founders to Dwnld's investors below.

Co-founders at Dwnld

AK
Alexandra KeatingCo-Founder & CEO
FL
Fritz LanmanCo-Founder & Executive Chairman

Director; Lead Independent Director (Greylock board partner) at Figma

Intro paths

β†’

In turn, John can intro founders to Figma's investors below.

Figma's investors Β· 33

137 VenturesSan FranciscoAlkeon InnovationNew YorkAltos VenturesBurlingame Β· $1M – $3M checksAmplify.LALos Angeles Β· $500K – $5M checksAndreessen HorowitzMenlo Park Β· $500K – $50M checks Β· since 2020Atlassian Ventures$1M – $5M checksBase PartnersSΓ£o PauloBase10 PartnersSan Francisco Β· $100K – $5M checksCoatue ManagementNew York Β· $1M – $20M checksDurable Capital PartnersBethesdaFelicis VenturesSan Francisco Β· $500K – $15M checksFidelity InvestmentsBostonFirst Round CapitalSan Francisco Β· $1M – $7M checksFounders FundSan Francisco Β· $500K – $5M checks Β· since 2020Fuel CapitalCovina Β· $500K – $1M checksGeodesic CapitalFoster CityGreylock PartnersSan Francisco Β· $100K – $200M checks Β· since 2014HaystackSan Francisco Β· $1M – $3M checksICONIQSan FranciscoIndex VenturesSan Francisco Β· $540K – $10.8M checks Β· since 2019IVP (Institutional Venture Partners)San Francisco Β· $15M – $75M checksKearny JacksonSan Francisco Β· $200K – $2M checksKleiner PerkinsMenlo Park Β· $500K – $15M checks Β· since 2024m]x[v CapitalNew YorkMosaic VenturesLondon Β· $1M – $10M checksNext Legacy PartnersPortola ValleyNext Play VenturesMenlo Park Β· $100K – $5M checksOutcast VenturesSan Francisco Β· $500K checksRitual CapitalMenlo Park Β· $100K – $500K checksSequoia CapitalMenlo Park Β· $4.4M – $214M checks Β· since 2019super{set}San FranciscoWestbound Equity PartnersPalo Alto Β· $750K – $2M checksWndrCoRedwood City Β· $100K – $10M checks

Co-founders at Figma

EW
Evan WallaceCo-Founder (former Chief Technology Officer)
DFDylan Field
Dylan FieldCo-founder & CEO

Experience

  1. Gigascale CapitalAdvising Partnercurrent2026 β€” present
  2. FigmaBoard Membercurrent2014 β€” present
  3. DuolingoBoard Membercurrent2021 β€” present
  4. VotingWorksBoard Membercurrent2018 β€” present
  5. NuroBoard Membercurrent2019 β€” present
  6. Greylock PartnersVenture Partnercurrent2019 β€” present
  7. Greylock PartnersPartner2011 β€” 2019
  8. Stanford UniversityConsulting Assistant Professor & Lecturercurrent2008 β€” present
  9. Code for AmericaChair, Board Of Directors2017 β€” 2025
  10. Code for AmericaBoard Member2012 β€” 2025
  11. ClockwiseBoard Member2018 β€” 2025
  12. CaffeineBoard Member2016 β€” 2024
  13. MozillaBoard Member2006 β€” 2014
  14. MozillaCEO2008 β€” 2010
  15. MozillaChief Operating Officer & Board Member2005 β€” 2007
  16. TripItBoard of Directors2008 β€” 2010
  17. Sunnyvale Public LibraryMember, Board of Trustees2005 β€” 2009
  18. Open Source Applications FoundationMember, Board of Directors2001 β€” 2008
  19. Reactivity (acquired by Cisco)CTO, VP Products, Board of Directors & Founder2001 β€” 2004
  20. Reactivity (acquired by Cisco)CEO, Board of Directors & Founder1998 β€” 2000
  21. CenterRunMember, Board of Directors2000 β€” 2003
  22. Apple ComputerSenior Scientist1997
  23. Trilogy SoftwareDirector of Design1995 β€” 1996

Skills & more

Open Source27SaaS7Enterprise Software19Security4Mobile Applications27Interaction Design10Venture Capital36Mobile Internet8Technology Management11Seed Capital2Data Visualization2User Experience42
+12 moreProduct ManagementBusiness DevelopmentUser Interface DesignScalabilityStart-upsUser InterfaceStrategic PartnershipsUser-centered DesignWeb DevelopmentEntrepreneurshipMobile DevicesLeadership

From John's professional profile.

Deals Β· 2

FigmaBoard seat

via Greylock Partners Β· filed Dec 2015 Β· $14M raise

DwnldBoard seat

via Greylock Partners Β· filed Sep 2015 Β· $17.1M raise

Attribution from SEC filings, press coverage, and firm rosters.

Other disclosed investments

Led

DropboxInstagramQuipTumblr

Board seats

DuolingoNuroQuip

Sources: the firm's website Β· SEC Form D filings.

Colleagues at Greylock Partners

Network2

Investors connected to John in public investor directories β€” a warm-intro map.

Frequently asked questions

Who is John Lilly?
John Lilly is an Advising Partner at Gigascale Capital with roughly 30 years of experience as a founder, CEO, and investor. He previously spent 8 years as a General Partner at Greylock and was CEO of Mozilla, where he oversaw Firefox's growth. Earlier he co-founded the enterprise security company Reactivity, later acquired by Cisco, and held roles at Apple and Trilogy Software. He is also a Lecturer at Stanford's Graduate School of Business and holds BS and MS degrees from Stanford.
Where does John Lilly work?
John Lilly is Partner at Greylock Partners and Board Member (Partner, Greylock Partners) at Dwnld and Director; Lead Independent Director (Greylock board partner) at Figma.
What does John Lilly invest in?
John Lilly invests in Cybersecurity, Enterprise Software.
What boards does John Lilly sit on?
John Lilly holds tracked board seats at Duolingo, Dwnld, Figma, Figma, Nuro, Quip.
What companies has John Lilly invested in?
Investments tracked here include Dropbox, Figma, Instagram, Quip, Tumblr.
Where did John Lilly go to school?
John Lilly studied at Stanford University (MS, Computer Science); Stanford University (BS, Computer Systems Engineering).
Does John Lilly lead rounds?
Yes β€” John Lilly has led 5 tracked investments.

Same school

Shared employers