/People

John Lilly

Investor

Advising Partner at Gigascale Capital · Consulting Assistant Professor & Lecturer at Stanford University · Venture Partner at Greylock Partners · Board Member / Board Partner (Greylock) at Clockwise · Board Member; Venture Partner, Greylock Partners; Advising Partner, Gigascale Capital at Nuro · +1 more

San Francisco Bay Area

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You →Chad · together at Trilogy Software (1995–1996) →John

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About

John Lilly is an Advising Partner at Gigascale Capital with roughly 30 years of experience as a founder, CEO, and investor. He previously spent 8 years as a General Partner at Greylock and was CEO of Mozilla, where he oversaw Firefox's growth. Earlier he co-founded the enterprise security company Reactivity, later acquired by Cisco, and held roles at Apple and Trilogy Software. He is also a Lecturer at Stanford's Graduate School of Business and holds BS and MS degrees from Stanford.

Likely emails· guessed from name + firm domain, not firm-published

Invests in

CybersecurityEnterprise Software

Board Member / Board Partner (Greylock) at Clockwise

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Co-founders at Clockwise

MMMatt Martin
GLGary Lerhaupt
Gary LerhauptFounder
MGMike Grinolds

Board Member; Venture Partner, Greylock Partners; Advising Partner, Gigascale Capital at Nuro

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Co-founders at Nuro

DFDave Ferguson
Dave FergusonFounder
JZJiajun Zhu
Jiajun ZhuFounder
DF
Dave Ferguson Jiajun ZhuFounder
DF
David FergusonFounder

Board Member (Partner, Greylock Partners) at Dwnld

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Co-founders at Dwnld

AKAlexandra Keating
Alexandra KeatingCo-Founder & CEO
FLFritz Lanman
Fritz LanmanCo-Founder & Executive Chairman

Experience

  1. Gigascale CapitalAdvising Partnercurrent2026 — present
  2. FigmaBoard Membercurrent2014 — present
  3. DuolingoBoard Membercurrent2021 — present
  4. VotingWorksBoard Membercurrent2018 — present
  5. NuroBoard Membercurrent2019 — present
  6. Greylock PartnersVenture Partnercurrent2019 — present
  7. Greylock PartnersPartner2011 — 2019
  8. Stanford UniversityConsulting Assistant Professor & Lecturercurrent2008 — present
  9. Code for AmericaChair, Board Of Directors2017 — 2025
  10. Code for AmericaBoard Member2012 — 2025
  11. ClockwiseBoard Member2018 — 2025
  12. CaffeineBoard Member2016 — 2024
  13. MozillaBoard Member2006 — 2014
  14. MozillaCEO2008 — 2010
  15. MozillaChief Operating Officer & Board Member2005 — 2007
  16. TripItBoard of Directors2008 — 2010
  17. Sunnyvale Public LibraryMember, Board of Trustees2005 — 2009
  18. Open Source Applications FoundationMember, Board of Directors2001 — 2008
  19. Reactivity (acquired by Cisco)CTO, VP Products, Board of Directors & Founder2001 — 2004
  20. Reactivity (acquired by Cisco)CEO, Board of Directors & Founder1998 — 2000
  21. CenterRunMember, Board of Directors2000 — 2003
  22. Apple ComputerSenior Scientist1997
  23. Trilogy SoftwareDirector of Design1995 — 1996

Skills & more

Security4Mobile Applications27Interaction Design10Venture Capital36Mobile Internet8Leadership23Technology Management11Seed Capital2Data Visualization2Open Source27User Experience42Product Management81
+8 moreBusiness DevelopmentUser Interface DesignScalabilityStart-upsUser InterfaceStrategic PartnershipsSaaSUser-centered Design

From John's professional profile.

Deals · 2

FigmaBoard seat

via Greylock Partners · filed Dec 2015 · $14M raise

DwnldBoard seat

via Greylock Partners · filed Sep 2015 · $17.1M raise

Attribution from SEC filings, press coverage, and firm rosters.

Other disclosed investments

Led

DropboxInstagramQuipTumblr

Board seats

DuolingoNuroQuip

Investor

AirtopApprenteArcolAsideBasedashCensusCitrus LaneClearSlideClockwiseCoScreenCrewDaffyDescriptDropboxFeltFluxGowallaHUBUCInstagramIron FishMarley MedicalMessageMeMomentoMoonsenseNuroNuroOzloPlayerZeroQuipRootlySkiffSocketSpark MobilesparksSpindleTENYXTerratonTumblrTumblr, Inc.UnblockedVerst

Sources: the firm's website · SEC Form D filings · curated public sources.

Colleagues at Gigascale Capital

Network2

Investors connected to John in public investor directories — a warm-intro map.

Frequently asked questions

Who is John Lilly?
John Lilly is an Advising Partner at Gigascale Capital with roughly 30 years of experience as a founder, CEO, and investor. He previously spent 8 years as a General Partner at Greylock and was CEO of Mozilla, where he oversaw Firefox's growth. Earlier he co-founded the enterprise security company Reactivity, later acquired by Cisco, and held roles at Apple and Trilogy Software. He is also a Lecturer at Stanford's Graduate School of Business and holds BS and MS degrees from Stanford.
Where does John Lilly work?
John Lilly is Advising Partner at Gigascale Capital and Board Member / Board Partner (Greylock) at Clockwise and Board Member; Venture Partner, Greylock Partners; Advising Partner, Gigascale Capital at Nuro and Board Member (Partner, Greylock Partners) at Dwnld.
What does John Lilly invest in?
John Lilly invests in Cybersecurity, Enterprise Software.
What boards does John Lilly sit on?
John Lilly holds tracked board seats at Duolingo, Dwnld, Figma, Figma, Nuro, Quip.
What companies has John Lilly invested in?
Investments tracked here include Dropbox, Figma, Instagram, Quip, Tumblr, Airtop, Apprente, Arcol, Aside, Basedash, Census, Citrus Lane, and 35 more.
Where did John Lilly go to school?
John Lilly studied at Stanford University (MS, Computer Science); Stanford University (BS, Computer Systems Engineering).
Does John Lilly lead rounds?
Yes — John Lilly has led 5 tracked investments.

Founder mafias

Same school

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